Females arrive after the nests are completed, coming in from deeper waters. At first the females appear to be chased away from the nest by the males, but after a considerable amount of chasing, the females head toward the nest instead of away from it. Once the female is in the nest, the pair swims in circles side by side, with the bellies of both fish touching. The male then releases milt and the female releases eggs. Females may spawn in more than one nest, and more than one female may use the same nest. Sometimes more than one female will spawn with a male simultaneously. Females produce 1,500 to 1,700 eggs, depending on their size and age.
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